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The emerging field of multispecies studies, grounded in passionate immersion in the lives of fungi, microorganisms, animals, plants, and others, is opening up novel ways of engaging with worlds around us. This issue brings together some of the leading scholars in this field to explore what is at stake—epistemologically, politically, ethically—for different forms of life caught up in diverse relationships of knowing and living together. The collection takes us into the worlds of sheep and shepherds; of stones, worms, salmon, and forest-devouring beetles; of viruses and their elephants; of seals, crows, and lava flows in Hawaii; and finally of frogs-as-pregnancy-tests and possible agents of pathogenic fungal spread. Each of the contributors explores what difference curious and careful attention to others might make in our efforts to inhabit and coconstitute flourishing worlds in these difficult times.